Within Clade 2, the sister group relationship of Harrimanella with the Styphelioideae + Vaccinioideae clade is well supported (Fig. 7). However, this relationship is only supported by molecular data and there are no discernable morphological synapomorphies for Clade 2. The Styphelioideae + Vaccinioideae clade is diagnosed by only a single homoplasious morphological feature—a persistent, non-withering calyx (char. #41), but the group is strongly supported by molecular characters.
Harrimanelloideae Kron & Judd, subfam. nov. – Type genus: Harrimanella Cov.
Latin Diagnosis: Harrimanelloideae foliis alternis et ± ericoidiis, pilis solum unicellularis, floribus 5-mero, solitariis et terminatibus, bracteis et bracteolis nullis, stylo brevi et crasso differt.

Mat-forming shrubs, stem lacking pericyclic sheath of fibers, with homogeneous pith, evergreen, with alternate, small, acicular, ericoid-like, petiolate leaves, revolute in bud. Leaf epidermal cells not lignified. Indumentum of only unicellular hairs. Inflorescence terminal, flowers solitary; bracts and bracteoles lacking; calyx articulated with pedicel. Flowers pentamerous, actinomorphic. Calyx lobes moderate-sized, ± deciduous; corolla sympetalous, ± campanulate, the lobes large; stamens 10, included, the filaments ± straight, smooth; anthers tetrasporangiate, inverting early in development, ± smooth, with a pair of dorso-apical spurs at junction with filament, lacking a fibrous endothecium, without disintegration tissue dorsally, dehiscing by terminal, slit-like pores. Pollen in tetrads, without viscin threads. Ovary 5-locular, with axile placentation, superior; style slightly impressed, short and stout; stigma truncate. Fruit a loculicidal capsule; seeds with moderately elongate testa cells; embryo with 2 cotyledons.
Distribution Map of Harrimanelloideae throughout the World
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